The species is thought to have arisen in the middle or late Pliocene.

It is believed to have gone extinct in the Late Pliocene.

It is dated approximately 2.6 million years ago, and is part of the late Pliocene.

In the late Pliocene they were also present in Africa.

It was active in the late Pliocene and early Pleistocene times.

It did not develop at all until the late Pliocene, but apparently developed very rapidly with the first continental glaciation.

This probably took place in the Late Pliocene, roughly 4-3 mya.

It lived in eastern Africa in the early to late Pliocene.

It is dated to the late Pliocene to the early Pleistocene (2-6 million years ago).

Geologically, this area is part of a structural downwarp that occurred during the late Pliocene.