Three are located on the facial side of the tooth, and one on the lingual side.

A third major cusp, the protocone, is present on the lingual side of both upper molars.

The valleys on the labial and lingual (inner) sides of the molars meet at the midlines.

An even smaller ridge is located below the ridge on the lingual side.

They also share the presence of furrows on the lingual side of the teeth only, but whether this is a derived feature is uncertain.

The fourth premolar (p4) also has a high central cusp; in addition, there are smaller roots before and behind it on the lingual side.

It is found on the distal lingual side of the tooth.

The labial side of the tooth is taller and less worn than the lingual side.

There are three roots: two at the labial and one at the lingual side.

The m1, m2, and usually m3 have a mesolophid, a crest corresponding to the mesoloph but located on the lingual side.