When it is time, the males migrate to the water.
This disproportion is due to the fact that more males migrate to the United States, were as females usually stay.
Many males remain close to hibernacula during the spring and summer rather than migrating long distances like females.
The males migrate first, and establish nest sites in strong current areas in rock crevices or on ledges.
The male will migrate to the breeding sites where he will begin calling his mate in intervals that can last up to thirty seconds.
The males become mobile and migrate through the soil searching for females with which to mate.
In order to breed, the males migrate to the females' habitat.
Prior to spawning, males and females migrate upstream where possible.
In almost every case, males migrate to adjacent groups.
The practice of transhumant sheepherding required that for several months each year, males migrate to the lowlands to pasture their animals.