No doubt he felt deeply and expressed himself strongly, but he was, in fact, not always the principal culprit.

Rather, the new findings indicate, prematurity is the principal culprit.

While Greenhouse gases are the principal culprit, other factors may be involved, as summarized in this diagram.

Inequality in the United States has indeed increased over the past 15 years, but the tax code was not the principal culprit.

The principal culprits in all this, the President insisted, were the Congressional Democrats, and especially their leaders.

The company's head, Martin Feinstein, pointed to the tax laws as a principal culprit.

The principal culprit for the capital's plight, ecologists and Government officials agree, is the rapidly growing number of cars.

"The financiers are really the principal culprits," he said.

The defendants disputed over who among them was the principal culprit and when they agreed to kill the victim.

Meanwhile the scandal was discussed in political circles, with Fazio singled out as the principal culprit and there were repeated calls for his resignation.