Operating system runs on a version of the Linux Kernel (computer science).
It tricks the program into thinking it is running on an earlier version of Windows.
It is run on a shortened version of the former Adelaide Grand Prix Circuit.
The game runs on a heavily modified version of the Quake engine known as DarkPlaces engine.
Iowa is running a second round of tests on an improved version of the system this winter.
The site runs on an updated version of the Moodle open source distance learning platform.
Each game is implemented as if running on a modernized version of the classic platform it represents.
The framework runs on a version of the Common Language Runtime that is optimized for gaming to provide a managed execution environment.
The game runs on a slightly modified version of the White Wolf Storyteller System.
Let's not forget that Java likes to keep older versions around, and programs (malware) can request to specifically run on an older version.