The subordinate male also forfeits to the dominant male all sexual relations with local females.

These females may form new groups with subordinate males.

Zamir, nine year old subordinate male, is one of the highest ranking males in the troop and has his eye on the top job.

The subordinate males eased forward, shoulder to shoulder, to stand beside the leader, ready to lower their horns.

"Don't you even have the balls to be angry that she slept with a subordinate male?"

A subordinate male in a reproductive unit may help care for an infant when it is six months old.

The alpha male is always easy to discern, but there are sometimes ambiguous rankings among subordinate males.

The subordinate male may spawn with the female and then retreat to a safe place.

Larger and more dominant males will sometimes try to remove a subordinate male while he is mating with a female.

Our zoo had a clan of six, two dominant females and four subordinate males.