If the reintroduced wolves remain in Yellowstone, scientists believe, a transformation in relationships among the ecosystem's wild inhabitants will take place.

Though wolf populations have increased in Ukraine, wolves remain unprotected there and can be hunted year-round by permit-holders.

"The wolf will remain with us," Qhorin said.

During pregnancy, female wolves remain in a den located away from the peripheral zone of their territories, where violent encounters with other packs are more likely.

Antoine, who believed the first whelp to have fallen on 4 October, surmised that but one wolf remained.

The wolves remained behind as the three of them began their descent to the lodge.

The two wolves remained, amber eyes regarding the quin'on'a with suspicion.

It is useless for the sheep to pass resolutions in favour of vegetarianism, while the wolf remains of a different opinion.

The dogs were gone, carried off by the plague, but the wolves remained in the high mountain country, ready to serve the Imp of Satan.

Now his sign was gone, but his wolf remained.